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During World War II, Arnim Zola was Hydra's top scientist, and an ally to the Red Skull. Throughout the war, he would often be caught up in conflicts between Hydra and the soldiers of the Allied forces, notably Captain America and Bucky Barnes. At some point long after the war's end, Zola was able to turn his conciousness into a computer algorithm, and upload it into an android body. Because of his unaging robotic body, Zola was able to survive to the present, while continuing to aid Hydra the whole time.